Dr. John Cullen received his undergraduate and veterinary training at the University of Pennsylvania. Following graduation, he completed a small animal medical internship at Angell Memorial Animal Hospital in Boston. After a few years in small animal practice, Dr. Cullen pursued an anatomic pathology residency and graduate studies at the University of California, Davis, where he says his enthusiasm for hepatic pathology began. Dr. Cullen has been at North Carolina State University in Raleigh for the last 37 years, where he divides his time between diagnostics, teaching and research. He has written and co-authored a number of manuscripts and several book chapters on liver diseases of animals and was recently named a Distinguished Member of the American College of Veterinary Pathologists, where he has served on the council and as President.